School of the Environment



Lisa Shipley

Associate Director/Professor



  • NATRS 310-Methods of Wildlife Ecology
  • NATRS 446-Wildlife Habitat Ecology
  • NATRS 595-Seminar in Natural Resource Sciences

Research Interests:

  • Foraging ecology of mammals, interactions between plants and herbivores, and ecology and management of wild ungulates
  • The influence of animal size, digestive morphology, and feeding strategy on the diet and habitats selected by herbivores (both free-ranging and captive herbivores, such as woodland
    caribou, blue duikers, porcupines, and pygmy rabbits, are used in these studies).

Areas of Expertise

Dr. Lisa A. Shipley is a wildlife ecologist in the Department of Natural Resource Sciences at Washington State University. Her research interests include foraging ecology and nutrition of mammals, interactions between plants and herbivores at different spatial scales, and ecology and management of wild ungulates. Both free-ranging and captive herbivores, such as mule deer, pronghorn, elk, and pygmy rabbits are used in these studies. Lisa directs the Wild Ungulate Facility and the Pygmy Rabbit Captive Breeding Facility at WSU. She is the faculty advisor of the WSU Student Chapter of The Wildlife Society.

Selected Publications

  • Shipley, L. A., E. M. Davis, L. A. Felicetti, S. McClean, and J. S. Forbey.  In revision. Mechanisms for eliminating monoterpenes of sageabrush by specialist and generalist rabbits.  Journal of Chemical Ecology.
  • Wagoner, S. J., L. A. Shipley, R. C. Cook, and L. Hardesty.  In revision. Effects of cattle grazing on the nutritional ecology of mule deer.  Journal of Wildlife Management.
  • Cook, R. C., J. C. Cook, D. J. Vales, B, K. Johnson, S. M. McCorquodale, L. A. Shipley,  R. A. Riggs,  L. L. Irwin, S. L. Murphie, B. L. Murphie, F. Geyer, P. B. Hall, R. D. Spencer, K. A. Schoenecker, D. A. Immell, D. H. Jackson, B. L. Tiller, P. J. Miller, and L. Schmitz.  In revision. A multi-regional evaluation of nutritional condition and reproduction in elk.  Wildlife Monographs.
  • Scarlata, C. D., B. A. Elias, J. R. Godwin, R. A. Powell, D. Shepherdson, L. A. Shipley and J. L. Brown.  In press.  Relationship between fecal hormone concentrations and reproductive success in captive pygmy rabbits (Brachylagus idahoensis).  Journal of Mammalogy.
  • Scarlata, C. D., B. A. Elias, J. R. Godwin, R. A. Powell, D. Shepherdson, L. A. Shipley and J. L. Brown 2011. Characterizing gonadal and adrenal activity by fecal steroid analysis in pygmy rabbits (Brachylagus idahoensis).  General and Comparative Endocrinology 171:373-380.
  • Tollefson, T. L., L. A. Shipley, W. L. Myers, and N. Dasgupta. 2011. Forage quality's influence on mule deer fawns.  Journal of Wildlife Management 75:919-928.
  • McCusker, S., L. A. Shipley, T. N. Tollefson, M. Griffin, and E. A. Koutsos. 2011.  Effects of starch and fibre in pelleted diets on nutritional status of mule deer (Odocoileus hemionus) fawns. Journal of Animal Physiology and Animal Nutrition 95: 489 - 498.
  • Dungan,J. D., L. A. Shipley, and R. G. Wright. 2010. Activity patterns, foraging ecology and summer range carrying capacity of moose in Rocky Mountain National Park, Colorado,  Alces 46: 71-87.
  • Tollefson, T. L., L. A. Shipley, W. L. Myers, D. H. Keisler, and N. Dasgupta. 2010.  The influence of summer and autumn nutrition on body condition and reproduction in lactating mule deer.  Journal of Wildlife Management 74:974-986.
  • Shipley, L. A.  2010.  Fifty years of food and foraging in moose: Lessons in ecology from a model herbivore.  Alces 46:1-13.
  • McClure, M. F., and L. A. Shipley.  2009. Animal response to nested self-similar patches: A test with woolly bears. Oikos 118:653-662.
  • Johnstone-Yellin, T. L., L. A. Shipley, W. L. Myers, and H. S. Robinson.  2009.  To twin or not to twin? Tradeoffs in litter size and fawn survival in mule deer.  Journal of Mammalogy 90:453-460.
  • Shipley, L. A., J. S. Forbey, and B. D. Moore. 2009.  Revisiting the dietary niche: when    is a mammalian herbivore a specialist?  Integrative and Comparative Biology 49:274-290.  
  • Kendrick, E., L. A. Shipley, A. E. Hagerman, and L. M. Kelly.  2009. Fruit and fibre: The nutritional value of figs for a small tropical ruminant, the blue duiker (Cephalophus monticola). Journal of African Ecology 47:556-566.
  • DeGabriel, J. L., B. D. Moore, L. A. Shipley, A. K. Krockenberger, I. R. Wallis, C. N. Johnson, and W. J. Foley.  2009. Inter-population differences in the tolerance of a marsupial folivore to plant secondary metabolites.  Oecologia 161:534-548.
  • Shipley, L. A. 2008. Pygmy rabbits in peril in the U.S.A. 
  • Thines, N.J., L.A. Shipley, J.H. Bassman, J.R. Slusser and W. Gao 2008. UV-B eefectson nutritional Chemistry of Plants and Responses of a Mammalian Herbivore. Oecologia 156: 125-135.
  • Shipley, L. A.  2007.  The influence of bite size on foraging at larger spatial and temporal scales by mammalian herbivores.  Oikos 116:1964-1974.
  • Robbins, C. T., J. K. Fortin, K. D. Rode, S. D. Farley, and L. A. Shipley.  2007.  Optimizing protein intake as a foraging strategy to maximize mass gain in an omnivore.  Oikos 116:1675-1682.
  • Cook, R. C., T. R. Stephenson, W. L. Myers, J. G. Cook, and L. A. Shipley.  2007. Validating predictive models of nutritional condition for mule deer.  Journal of Wildlife Management 71:1934-1943.
  • Thines, N. J., L. A. Shipley, J. H. Bassman, J. K. Fellman, D. S. Mattison, and J. R. Slusser. 2007. Effects of enhanced UV-B radiation on plant chemistry: Nutritional consequences for a specialist and generalist lagomorph.Journal of Chemical Ecology 33:1025-1039.


School of the Environment
Washington State University
425 Heald Hall
Pullman, WA, 99164-2812

(509) 335-9182


Related Links:

School of the Environment, PO Box 642812, Washington State University, Pullman WA 99164-2812, 509-335-3009, Contact Us